T.  Sato, Kushiro Shitsugen Nature Guide, Siberian Salamander. Kushiro Onnenai: Japanese Society for Preservation of Birds, 1993.

M.  Matsui, Yoshikawa, N., Tominaga, A., Sato, T., Takenaka, S., Tanabe, S., Nishikawa, K., and Nakabayashi, S., ?Phylogenetic relationships of two Salamandrella species as revealed by mitochondrial DNA and allozyme variation (Amphibia: Caudata: Hynobiidae)?, Molecular Phylogenetics and Evolution, vol. 48, pp. 84 - 93, 2008.